Pre-Republic
The very early history of Coruscant is a bit sketchy and is not well known. Coruscant was considered by many to be the Human homeworld. Millennia ago, the Architects could have removed Humans from Coruscant with which to populate Corellia.
Around 200,000 BBY, the near-Human Taungs attempted to conquer the 13 baseline Human nations of the Battalions of Zhell. A volcano decimated the Zhell, the ash filling the skies for two years, so the Taungs adopted the name Dha Werda Verda (Warriors of Shadow) for themselves. The Human Zhell eventually recovered and drove the Taungs offworld - the Taungs fled to Roon.
One hundred millennia later, Coruscant was surveyed by the Columi, who dismissed the planet as a primitive disappointment, despite the already planet-spanning ecumenopolis of Galactic City. New buildings were built on the old. As a result, there was virtually no exposed land. In the forgotten underlevels of the city, there was darkness, pollution and crime. Higher up, there were government offices and penthouses owned by the elite. The lower fifty levels of the ecumenopolis is said to have last seen sunlight around 95,000 BBY.
Coruscant was one of many worlds conquered by the Infinite Empire of the Rakata, who used Human slaves to build the Star Forge in 30,000 BBY. Under Rakatan domination, the Humans of Coruscant's colonization attempts were limited to sleeper ships, which ended up on Alderaan, in the Tion Cluster, Seoul 5, Kuat, Alsakan, Axum, Anaxes, Atrisia, Metellos, Corulag, and many other worlds. The Rakata were eventually decimated by a massive plague, leading to slave revolutions on Coruscant and other subjected worlds.
Over the next two centuries, Coruscant was linked to other Core Worlds, including Corellia and Duro, by hyperspace cannons. It was during this time that the Coruscant government peacefully absorbed the nearby Azure Imperium. During these pre-Republic years, the languages of Coruscant and its neighbors meshed to become Old Galactic Standard.
Early Republic
Galactic City c. 5,000 BBY.
In 25,000 BBY, the Corellians and Duros invented the hyperdrive proper, allowing Coruscant to become the capital of a democratic union - the Galactic Republic. The planet remained the Republic's capital for twenty five thousand years. Shortly after the formation of the Republic, the Perlemian Trade Route was mapped, linking Coruscant to Ossus and bringing the Jedi Knights into the Republic. Over the next millennium, the Corellian Run was mapped, linking Coruscant to Corellia and beyond. Blasters were also invented on Coruscant around this time, and the famous Galactic Museum was constructed in 12,000 BBY.
From the very beginning, Coruscant, as the Republic's capital, was the primary objective in several wars. The earliest among these was the Tionese War with the Honorable Union of Desevro and Tion in 24,000 BBY, in which Coruscant was bombarded with Tionese pressure bombs. Other early battles included the Alsakan Conflicts, the Duinuogwuin Contention, the Great Hyperspace War, the Third Great Schism, the Great Droid Revolution, and the Great Sith War.
Following the devastation of Ossus, the Jedi Council took up residence in the Jedi Temple on Coruscant, to which many Jedi relics from Ossus were taken. The Temple was greatly expanded, including the building of the original Jedi Council Chamber. The Temple was repeatedly expanded including in 3,519 BBY, 2,519 BBY (when the Jedi Archives were built), 2,019 BBY, and 1,000 BBY (when the Temple spires were built).
In 2,973 BBY, the Senate Rotunda was founded, replacing the previous assembly place.
End of the Republic
As Darth Sidious gained in power (and made Anakin Skywalker Darth Vader in the process), and other major politicians and entities became more aggressive, political instability surged both in the Galaxy and on Coruscant, threatening Coruscant itself and inflicting much violence on and near Coruscant.
In 27 BBY, Coruscant was almost destroyed by Ashaar Khorda using the Infant of Shaa. Jango Fett, Zam Wesell, and Yarael Poof saved Coruscant, although Poof sacrificed himself to do so.
In 23 BBY, near the end of the Separatist Crisis, the Confederacy of Independent Systems began to become a significant threat against the Republic, causing several systems to secede, including Ando and Sy Myrth, resulting in a mass exodus of Aqualish and Sy Myrthians from Coruscant.
The Jedi Temple on Coruscant at sunset.
During these times, several assassination attempts were made on Padmé Amidala while she was on Coruscant, including one where her personal ship was destroyed and several of her handmaidens killed. Anakin Skywalker and Obi-Wan Kenobi were assigned to protect her. Eventually, they managed to foil an attempt at Padmé's penthouse by Zam Wesell, resulting in an airspeeder chase through Coruscant, ending in a showdown at the Outlander Club in the lower levels. The Jedi caught Zam, but she was killed by a mysterious armored figure, leading Obi-Wan to Kamino.
During most of the Clone Wars, Coruscant was subject to frequent Separatist terrorist acts and assassinations of leading Republic figures. These included the holding of Senator Meena Tills hostage by Korunnai terrorists in 22 BBY (actually cover for an attempt to turn the Corporate Sector against the Republic). 21 BBY saw a wave of such disasters, including the Coruscant Insurrection and the assassinations of Finis Valorum and Senator Viento.
Such incidents prompted the Republic to implement numerous changes on Coruscant in the name of increased security. Huge stretches of the cityscape were transformed into military staging areas, and clone troopers were seen everywhere. The fanatical pro-Human COMPOR was founded, the local SAGroup led by Nenevanth Tion marching in a patriotism parade. COMPOR pressured Tannon Praji into deporting from the capital all members of species whose homeworld had joined the Confederacy. Vice-Admiral Terrinald Screed was placed in charge of Coruscant Planetary Defense.
Yet in 19 BBY, the Confederacy of Independent Systems succeeded in making a raid on Coruscant. Using secret hyperspace routes through the Deep Core provided by Darth Sidious, a Separatist fleet under the command of the dreaded General Grievous made a raid on the city planet. While the Separatist fleet engaged the Republic fleet, an army of battle droids landed on Galactic City. The invasion distracted the defending Jedi and clone forces while General Grievous and a squad of his elite MagnaGuards captured the Supreme Chancellor Palpatine. However, Anakin Skywalker and Obi-Wan Kenobi succeeded in rescuing the Supreme Chancellor from the infamous Separatist flagship Invisible Hand. The Battle of Coruscant eventually turned to the Republic's favor and the Separatist's armada was forced into a full retreat.
However, the battle had shaken much of the population, and large portions of the city had been decimated by falling ships or collateral damage caused in the vicious fighting.
Near the end of the Clone Wars, Coruscant was transformed into Imperial Center following Chancellor Palpatine's Declaration of a New Order, reorganizing the Galactic Republic into the Galactic Empire.
Galactic Empire
The skyscrapers of Coruscant.
While the planet's name was officially changed during Palpatine's rule, most denizens of the Empire continued to call the planet-city by its old name. As part of the Humanocentric policies of the galaxy, nonhuman species were also compelled to live in segregated parts (also known as "ethnic neighborhoods") of Imperial City, such as Invisec. Those that did enter other parts, whether by intention or mistake, were punished brutally.
During Imperial times, a proclamation by Palpatine made Imperial Center the only planet in the galaxy where political science students could do postgraduate work. The reason was twofold: foremost was the hope that the future politicians and technocrats of the galaxy would be more likely to accept the values and rule of the New Order; second was that the brightest of them could be detained, either voluntarily or by force, on Center to assist in the running of the Empire.
Emperor Palpatine also began to shape Coruscant with his own designs, and this included the installment of thousands of surveillance systems throughout the ecumenopolis. Massive shield generators were placed all over Coruscant, and where they overlapped, powerful storms took place, giving the planet a gothic look, which mirrored the utter desperation of many of its inhabitants as they choked under Palpatine's iron rule.
The Emperor also had the Palace of the Republic rebuilt and renamed the Imperial Palace. The Imperial Palace was a massive pyramidal structure and was the largest construction project in Coruscant. Other distinctive structures of the Galactic Republic, such as the Galactic Senate and the Jedi Temple, were either destroyed or eclipsed by newer buildings.